Heptapteridae

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Heptapteridae
Rhamdia quelen
Scientific classification
Kingdom: Animalia Phylum: Chordata Class: Actinopterygii Order: Siluriformes Superfamily: Pimelodoidea Family: Heptapteridae Gill, 1861[1]
Genera
Acentronichthys Brachyglanis Brachyrhamdia Cetopsorhamdia Chasmocranus Gladioglanis Goeldiella Heptapterus Horiomyzon Imparfinis Leptorhamdia Mastiglanis Myoglanis Nannoglanis Nemuroglanis Pariolius Phenacorhamdia Pimelodella Rhamdella Rhamdia Rhamdioglanis Rhamdiopsis Taunayia

Heptapteridae is a family of catfish that originate from Mexico to South America.^[2] Heptapteridae is derived from Greek, hepta meaning seven and pteron meaning fin.^[3]

The diversity of this family is poorly known and there are many species yet to be described.^[2] So far some 190 species have been described. This family is equivalent to the previously recognized Rhamdiinae, a subfamily of the family Pimelodidae.^[2] However, molecular evidence shows that this family is a part of the superfamily Pimelodoidea along with Pimelodidae, Pseudopimelodidae, and Conorhynchos.^[4]

The skin of these fish is usually naked (scaleless). They exhibit three pairs of barbels. They have a large adipose fin, and their caudal fin is deeply forked. However, there are no external characteristics that are unique to this family that allow it to be differentiated from Pimelodidae.^[2]

Heptapteridae includes a few troglobitic species in the genera Pimelodella, Rhamdia, and Taunayia, as well as in an undescribed genus.^[5]

[edit] Species

There are 189 species in 24 genera:^[6]

• Genus Acentronichthys Eigenmann & Eigenmann, 1889.
  • Acentronichthys leptos Eigenmann & Eigenmann, 1889.
• Genus Brachyglanis Eigenmann, 1912.
  • Brachyglanis frenatus Eigenmann, 1912.
  • Brachyglanis magoi Fernández-Yépez, 1967.
  • Brachyglanis melas Eigenmann, 1912.
  • Brachyglanis microphthalmus Bizerril, 1991.
  • Brachyglanis nocturnus Myers, 1928.
  • Brachyglanis phalacra Eigenmann, 1912.
• Genus Brachyrhamdia Myers, 1927.
  • Brachyrhamdia heteropleura (Eigenmann, 1912).
  • Brachyrhamdia imitator Myers, 1927.
  • Brachyrhamdia marthae Sands & Black, 1985.
  • Brachyrhamdia meesi Sands & Black, 1985.
  • Brachyrhamdia rambarrani (Axelrod & Burgess, 1987).
• Genus Cetopsorhamdia Eigenmann & Fisher, 1916.
  • Cetopsorhamdia boquillae Eigenmann & Fisher, 1922.
  • Cetopsorhamdia filamentosa Fowler, 1945.
  • Cetopsorhamdia iheringi Schubart & Gomes, 1959.
  • Cetopsorhamdia insidiosa (Steindachner, 1915).
  • Cetopsorhamdia molinae Miles, 1943.
  • Cetopsorhamdia nasus Eigenmann & Fisher, 1916.
  • Cetopsorhamdia orinoco Schultz, 1944.
  • Cetopsorhamdia phantasia Stewart, 1985.
  • Cetopsorhamdia picklei Schultz, 1944.
• Genus Chasmocranus Eigenmann, 1912.
  • Chasmocranus brachynemus Gomes & Schubart, 1958.
  • Chasmocranus brevior Eigenmann, 1912.
  • Chasmocranus chimantanus Inger, 1956.
  • Chasmocranus longior Eigenmann, 1912.
  • Chasmocranus lopezi Miranda Ribeiro, 1968.
  • Chasmocranus peruanus Eigenmann & Pearson, 1942.
  • Chasmocranus quadrizonatus Pearson, 1937.
  • Chasmocranus rosae Eigenmann, 1919.
  • Chasmocranus surinamensis (Bleeker, 1862).
  • Chasmocranus truncatorostris Borodin, 1927.
• Genus Gladioglanis Ferraris & Mago-Leccia, 1989.
  • Gladioglanis conquistador Lundberg, Bornbusch & Mago-Leccia, 1991.
  • Gladioglanis machadoi Ferraris & Mago-Leccia, 1989.
• Genus Goeldiella Eigenmann & Norris, 1900.
  • Goeldiella eques (Müller & Troschel, 1848).
• Genus Heptapterus Bleeker, 1858.
  • Heptapterus bleekeri Boeseman, 1953.
  • Heptapterus fissipinnis Miranda Ribeiro, 1911.
  • Heptapterus multiradiatus Ihering, 1907.
  • Heptapterus mustelinus (Valenciennes, 1835).
  • Heptapterus stewarti Haseman, 1911.
  • Heptapterus sympterygium Buckup, 1988.
  • Heptapterus tapanahoniensis Mees, 1967.
  • Heptapterus tenuis Mees, 1986.
• Genus Horiomyzon Stewart, 1986.
  • Horiomyzon retropinnatus Stewart, 1986.
• Genus Imparfinis Eigenmann & Norris, 1900.
  • Imparfinis borodini Mees & Cala, 1989.
  • Imparfinis cochabambae (Fowler, 1940).
  • Imparfinis guttatus (Pearson, 1924).
  • Imparfinis hasemani Steindachner, 1915.
  • Imparfinis hollandi Haseman, 1911.
  • Imparfinis lineatus (Bussing, 1970).
  • Imparfinis longicauda (Boulenger, 1887).
  • Imparfinis microps Eigenmann & Fisher, 1916.
  • Imparfinis minutus (Lütken, 1874).
  • Imparfinis mirini Haseman, 1911.
  • Imparfinis mishky Almiron, Casciotta, Bechara et al. 2007.^[7]
  • Imparfinis nemacheir (Eigenmann & Fisher, 1916).
  • Imparfinis pijpersi (Hoedeman, 1961).
  • Imparfinis piperatus Eigenmann & Norris, 1900.
  • Imparfinis pristos Mees & Cala, 1989.
  • Imparfinis pseudonemacheir Mees & Cala, 1989.
  • Imparfinis schubarti (Gomes, 1956).
  • Imparfinis spurrellii (Regan, 1913).
  • Imparfinis stictonotus (Fowler, 1940).
• Genus Leptorhamdia Eigenmann, 1918.
  • Leptorhamdia essequibensis (Eigenmann, 1912).
  • Leptorhamdia marmorata Myers, 1928.
  • Leptorhamdia schultzi (Miranda Ribeiro, 1964).
• Genus Mastiglanis Bockmann, 1994.
  • Mastiglanis asopos Bockmann, 1994.
• Genus Myoglanis Eigenmann, 1912.
  • Myoglanis aspredinoides Do Nascimiento & Lundberg, 2005.
  • Myoglanis koepckei Chang, 1999.
  • Myoglanis potaroensis Eigenmann, 1912.
• Genus Nannoglanis Boulenger, 1887.
  • Nannoglanis fasciatus Boulenger, 1887.
• Genus Nemuroglanis Eigenmann & Eigenmann, 1889.
  • Nemuroglanis lanceolatus Eigenmann & Eigenmann, 1889.
  • Nemuroglanis mariai (Schultz, 1944).
  • Nemuroglanis pauciradiatus Ferraris, 1988.
• Genus Pariolius Cope, 1872.
  • Pariolius armillatus Cope, 1872.
• Genus Phenacorhamdia Dahl, 1961.
  • Phenacorhamdia anisura (Mees, 1987).
  • Phenacorhamdia boliviana (Pearson, 1924).
  • Phenacorhamdia hoehnei (Miranda Ribeiro, 1914).
  • Phenacorhamdia macarenensis Dahl, 1961.
  • Phenacorhamdia nigrolineata Zarske, 1998.
  • Phenacorhamdia somnians (Mees, 1974).
  • Phenacorhamdia tenebrosa (Schubart, 1964).
  • Phenacorhamdia unifasciata Britski, 1993.
• Genus Pimelodella Eigenmann & Eigenmann, 1888.
  • Pimelodella altipinnis (Steindachner, 1864).
  • Pimelodella australis Eigenmann, 1917.
  • Pimelodella avanhandavae Eigenmann, 1917.
  • Pimelodella boliviana Eigenmann, 1917.
  • Pimelodella boschmai Van der Stigchel, 1964.
  • Pimelodella brasiliensis (Steindachner, 1877).
  • Pimelodella breviceps (Kner, 1857).
  • Pimelodella buckleyi (Boulenger, 1887).
  • Pimelodella chagresi (Steindachner, 1876).
  • Pimelodella chaparae Fowler, 1940.
  • Pimelodella conquetaensis Ahl, 1925.
  • Pimelodella cristata (Müller & Troschel, 1848).
  • Pimelodella cruxenti Fernández-Yépez, 1950.
  • Pimelodella cyanostigma (Cope, 1870).
  • Pimelodella dorseyi Fowler, 1941.
  • Pimelodella eigenmanni (Boulenger, 1891).
  • Pimelodella eigenmanniorum (Miranda Ribeiro, 1911).
  • Pimelodella elongata (Günther, 1860).
  • Pimelodella enochi Fowler, 1941.
  • Pimelodella eutaenia Regan, 1913.
  • Pimelodella figueroai Dahl, 1961.
  • Pimelodella geryi Hoedeman, 1961.
  • Pimelodella gracilis (Valenciennes, 1835).
  • Pimelodella griffini Eigenmann, 1917.
  • Pimelodella grisea (Regan, 1903).
  • Pimelodella hartii (Steindachner, 1877).
  • Pimelodella hartwelli Fowler, 1940.
  • Pimelodella hasemani Eigenmann, 1917.
  • Pimelodella howesi Fowler, 1940.
  • Pimelodella itapicuruensis Eigenmann, 1917.
  • Pimelodella kronei (Miranda Ribeiro, 1907).
  • Pimelodella lateristriga (Lichtenstein, 1823).
  • Pimelodella laticeps Eigenmann, 1917.
  • Pimelodella laurenti Fowler, 1941.
  • Pimelodella linami Schultz, 1944.
  • Pimelodella macrocephala (Miles, 1943).
  • Pimelodella macturki Eigenmann, 1912.
  • Pimelodella martinezi Fernández-Yépez, 1970.
  • Pimelodella meeki Eigenmann, 1910.
  • Pimelodella megalops Eigenmann, 1912.
  • Pimelodella megalura Miranda Ribeiro, 1918.
  • Pimelodella metae Eigenmann, 1917.
  • Pimelodella modesta (Günther, 1860).
  • Pimelodella montana Allen, 1942.
  • Pimelodella mucosa Eigenmann & Ward, 1907.
  • Pimelodella nigrofasciata (Perugia, 1897).
  • Pimelodella notomelas Eigenmann, 1917.
  • Pimelodella odynea Schultz, 1944.
  • Pimelodella ophthalmica (Cope, 1878).
  • Pimelodella pallida Dahl, 1961.
  • Pimelodella pappenheimi Ahl, 1925.
  • Pimelodella parnahybae Fowler, 1941.
  • Pimelodella parva Güntert, 1942.
  • Pimelodella pectinifer Eigenmann & Eigenmann, 1888.
  • Pimelodella peruana Eigenmann & Myers, 1942.
  • Pimelodella peruensis Fowler, 1915.
  • Pimelodella procera Mees, 1983.
  • Pimelodella reyesi Dahl, 1964.
  • Pimelodella roccae Eigenmann, 1917.
  • Pimelodella rudolphi Miranda Ribeiro, 1918.
  • Pimelodella serrata Eigenmann, 1917.
  • Pimelodella spelaea Trajano, Reis & Bichuette, 2004.
  • Pimelodella steindachneri Eigenmann, 1917.
  • Pimelodella taeniophora (Regan, 1903).
  • Pimelodella taenioptera Miranda Ribeiro, 1914.
  • Pimelodella tapatapae Eigenmann, 1920.
  • Pimelodella transitoria Miranda Ribeiro, 1907.
  • Pimelodella vittata (Lütken, 1874).
  • Pimelodella wesselii (Steindachner, 1877).
  • Pimelodella witmeri Fowler, 1941.
  • Pimelodella yuncensis .
• Genus Rhamdella Eigenmann & Eigenmann, 1888.
  • Rhamdella aymarae Miquelarena & Menni, 1999.
  • Rhamdella cainguae Bockmann and Miquelarena, 2008.
  • Rhamdella eriarcha (Eigenmann & Eigenmann, 1888).
  • Rhamdella exsudans (Jenyns, 1842).
  • Rhamdella gilli (Starks, 1906).
  • Rhamdella ignobilis Steindachner, 1907.
  • Rhamdella jenynsii (Günther, 1864).
  • Rhamdella leptosoma Fowler, 1914.
  • Rhamdella longipinnis Borodin, 1927.
  • Rhamdella longiuscula Lucena & da Silva, 1991.
  • Rhamdella montana Eigenmann, 1913.
  • Rhamdella papariae Fowler, 1941.
  • Rhamdella robinsoni Fowler, 1941.
  • Rhamdella rusbyi Pearson, 1924.
  • Rhamdella wolfi Fowler, 1941.
• Genus Rhamdia Bleeker, 1858.
  • Rhamdia enfurnada Bichuette & Trajano, 2005.
  • Rhamdia foina (Müller & Troschel, 1848).
  • Rhamdia guasarensis Do Nascimiento, Provenzano & Lundberg, 2004.
  • Rhamdia humilis (Günther, 1864).
  • Rhamdia itacaiunas Silfvergrip, 1996.
  • Rhamdia jequitinhonha Silfvergrip, 1996.
  • Rhamdia laluchensis Weber, Allegrucci & Sbordoni, 2003.
  • Rhamdia laticauda (Kner, 1857).
  • Rhamdia laukidi Bleeker, 1858.
  • Rhamdia macuspanensis Weber & Wilkens, 1998.
  • Rhamdia muelleri (Günther, 1864).
  • Rhamdia nicaraguensis (Günther, 1864).
  • Rhamdia parryi Eigenmann & Eigenmann, 1888.
  • Rhamdia poeyi Eigenmann & Eigenmann, 1888.
  • Rhamdia quelen (Quoy & Gaimard, 1824).
  • Rhamdia reddelli Miller, 1984.
  • Rhamdia xetequepeque Silfvergrip, 1996.
• Genus Rhamdioglanis Ihering, 1907.
  • Rhamdioglanis frenatus Ihering, 1907.
  • Rhamdioglanis transfasciatus Miranda Ribeiro, 1908.
• Genus Rhamdiopsis Haseman, 1911.
  • Rhamdiopsis microcephala (Reinhardt, 1874).
  • Rhamdiopsis moreirai Haseman, 1911.
• Genus Taunayia Miranda Ribeiro, 1918.
  • Taunayia bifasciata (Eigenmann & Norris, 1900).

[edit] References

1. ^ Heptapteridae (TSN 679381). Integrated Taxonomic Information System. Retrieved on 07 April 2007.
2. ^ ^{a b c d} Nelson, Joseph, S. (2006). Fishes of the World. John Wiley & Sons, Inc.. ISBN 0471250317. 
3. ^ "Heptapteridae". FishBase. Ed. Rainer Froese and Daniel Pauly. Apr 2007 version. N.p.: FishBase, 2007.
4. ^ Sullivan, JP; Lundberg JG; Hardman M (2006). "A phylogenetic analysis of the major groups of catfishes (Teleostei: Siluriformes) using rag1 and rag2 nuclear gene sequences". Mol Phylogenet Evol. 41 (3): 636–62. doi:10.1016/j.ympev.2006.05.044. 
5. ^ Bichuette, Maria Elina; Trajano, Eleonora (2005). "A new cave species of Rhamdia (Siluriformes: Heptapteridae) from Serra do Ramalho, northeastern Brazil, with notes on ecology and behavior". Neotropical Ichthyology 3 (4): 587–595. 
6. ^ Ferraris, Carl J., Jr. (2007). "Checklist of catfishes, recent and fossil (Osteichthyes: Siluriformes), and catalogue of siluriform primary types" (PDF). Zootaxa 1418: 180–202. http://mapress.com/zootaxa/2007f/zt01418p300.pdf. 
7. ^ Almiron et al. (2007). "Imparfinis mishky (Siluriformes, Heptapteridae) a new species from the rios Parana and Uruguay basins in Argentina.". Revue Suisse de Zoologie 114 (4): 817–824.